The length of the action is the difference. The short action is based on supporting at a maximum the overall length of the .308 Winchester cartrige and it will support all cartriges of that length and shorter. The heavier and longer "long action" supports cartriges the length of the .30-06. There is another classification as well and that is the magnum action length based on cartiges the length of the .375 H&H and above. Each larger action is bigger and heavier to carry as well as being longer. Therefore many hunters prefer the small action size so that their rifle can be lighter to carry. If they choose a magnum length, they get a very heavy rifle to carry.
.223 .308 22-250 .243 examples normally used in short action, most short action ammo can also be used in a long action, long action examples 30/06 .300 win mag 7mm rem mag .270 winchester. long action ammo could be loaded one at a time in a short action but will be to long for the magazine.
I'm a little confused by this.... are you saying rifles can be fired with cartridges other than the rounds for which they are chambered?
Ie, 30-06 in 308 and or vice versa?
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